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doi: 10.1164/rccm.201005-0756OC 72 HaworthCSFowerakerJEWilkinsonPKenyonRFBiltonD

Given that the main structural difference between these two enzymes is the presence of a CBM loop insert in Bd GUS, the CBM loop may explain this difference in activity, as it may occlude a substrate like p NPG from being properly positioned in the active site

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